READING COMPREHENSION
Finish-born botanist William Nylander taught at the University of Helsinki for a number of years and later moved to Paris, where he lived until his death at the end of the nineteenth century. During the second half of the last century, he became a prominent figure in the field of lichenology. Botanists from all over the world sent samples to his laboratory to be analyzed and classified. It can be said without exaggeration that four of five lichens bear his name.
He was the first to realize the importance of using chemical reagents in the taxonomy of lichens. He selected the most common reagents used by the chemists of his time. Lichenologists all over the world still use these reagents, including tincture of iodine and hypochlorite in their laboratories. During the first half of the nineteenth century, a Japanese named Abahima added only one chemical product- Phenol diamines. Nylander was also responsible for discovering that the atmosphere of big cities hindered the lichens’ development and cause them to disappear. Now they are used to detect atmospheric pollution. Nevertheless, he considered lichens to be simple plants and vehemently opposed the widely accepted modern theories that lichens are compound species formed by two discordant elements: algae and fungi.

According to the nineteenth-century theories, which elements from the composition of lichens?

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